- Used in the "nuclear shuffle" similar to that found in croziers during sexual reproduction.
- Ensures that two adjacent hyphal cells (divided by septa) each have 2 different nuclei from mating with hyphae of another sexual type.
- C.J. Alexopolous, Charles W. Mims, M. Blackwell et al., Introductory Mycology, 4th ed. (John Wiley and Sons, Hoboken NJ, 2004) ISBN 0-471-52229-5