|Common Chimpanzeein Cameroon's South Province|
|distribution of Pan spp.|
distribution of Pan spp.
Chimpanzee, often shortened to chimp, is the common name for the two extant species of apes in the genus Pan. The better known chimpanzee is Pan troglodytes, the Common Chimpanzee, living primarily in West, and Central Africa. Its cousin, the Bonobo or "Pygmy Chimpanzee" as it is known archaically, Pan paniscus, is found in the forests of the Democratic Republic of the Congo. The Congo River forms the boundary between the two species. Chimpanzees are members of the Hominidae family, along with gorillas, humans, and orangutans, and the two chimpanzee species are the closest living relatives to humans.
A fully grown adult male chimpanzee can weigh from 35-70 kilograms (75-155 lb) and stand 0.9-1.2 metres (3-4 ft) tall, while females usually weigh 26-50 kg (57-110 lb) and stand 0.66-1 m (2-3½ ft) tall.
Chimpanzees rarely live past the age of 40 in the wild, but have been known to reach the age of more than 60 in captivity. Cheeta, star of Tarzan is still alive as of 2008 at the age of 76, making him the oldest known chimpanzee in the world.
Anatomical differences between the Common Chimpanzee and the Bonobo are slight, but in sexual and social behaviour there are marked differences. Common Chimpanzees have an omnivorous diet, a troop hunting culture based on beta males led by an alpha male, and highly complex social relationships. Bonobos, on the other hand, have a mostly frugivorous diet and an egalitarian, nonviolent, matriarchal, sexually receptive behaviour. The exposed skin of the face, hands and feet varies from pink to very dark in both species, but is generally lighter in younger individuals, darkening as maturity is reached. Bonobos have proportionately longer upper limbs and tend to walk upright more often than the Common Chimpanzee. A University of Chicago Medical Centre study has found significant genetic differences between chimpanzee populations. Different groups of Chimpanzees also have different cultural behaviour with preferences for types of tools.
History of human interaction
Africans have had contact with chimpanzees for millennia. Chimpanzees have been kept as domesticated pets for centuries in a few African villages, especially in Congo. The first recorded contact of Europeans with chimps took place in present-day Angola during the 1600s. The diary of Portuguese explorer Duarte Pacheco Pereira (1506), preserved in the Portuguese National Archive (Torre do Tombo), is probably the first European document to acknowledge that chimpanzees built their own rudimentary tools.
The first use of the name "chimpanzee", however, did not occur until 1738. The name is derived from a Tshiluba language term "kivili-chimpenze", which is the local name for the animal and translates loosely as "mockman" or possibly just "ape". The colloquialism "chimp" was most likely coined some time in the late 1870s. Biologists applied Pan as the genus name of the animal. Chimps as well as other apes had also been purported to have been known to Western writers in ancient times, but mainly as myths and legends on the edge of Euro-Arabic societal consciousness, mainly through fragmented and sketchy accounts of European adventurers. Apes are mentioned variously by Aristotle, as well as the Bible, where apes and baboons are described as having been collected by Solomon in 1 Kings 10:22.
When chimpanzees first began arriving on the European continent, European scientists noted the inaccuracy of some ancient descriptions, which often reported that chimpanzees had horns and hooves. The first of these early transcontinental chimpanzees came from Angola and were presented as a gift to Frederick Henry, Prince of Orange in 1640, and were followed by a few of its brethren over the next several years. Scientists who examined these rare specimens were baffled, and described these first chimpanzees as "pygmies", and noted the animals' distinct similarities to humans. The next two decades would see a number of the creatures imported into Europe, mainly acquired by various zoological gardens as entertainment for visitors.
Darwin's theory of evolution (published in 1859) spurred scientific interest in chimpanzees, as in much of life science, leading eventually to numerous studies of the animals in the wild and captivity. The observers of chimpanzees at the time were mainly interested in behaviour as it related to that of humans. This was less strictly and disinterestedly scientific than it might sound, with much attention being focused on whether or not the animals had traits that could be considered 'good'; the intelligence of chimpanzees was often significantly exaggerated. At one point there was even a scheme drawn up to domesticate chimpanzees in order to have them perform various menial tasks (i.e. factory work). By the end of the 1800s chimpanzees remained very much a mystery to humans, with very little factual scientific information available.
The 20th century saw a new age of scientific research into chimpanzee behaviour. Before 1960, almost nothing was known about chimpanzee behaviour in their natural habitat. In July of that year, Jane Goodall set out to Tanzania's Gombe forest to live among the chimpanzees. Her discovery that chimpanzees made and used tools was groundbreaking, as humans were previously believed to be the only species to do so. The most progressive early studies on chimpanzees were spearheaded primarily by Wolfgang Köhler and Robert Yerkes, both of whom were renowned psychologists. Both men and their colleagues established laboratory studies of chimpanzees focused specifically on learning about the intellectual abilities of chimpanzees, particularly problem-solving. This typically involved basic, practical tests on laboratory chimpanzees, which required a fairly high intellectual capacity (such as how to solve the problem of acquiring an out-of-reach banana). Notably, Yerkes also made extensive observations of chimpanzees in the wild which added tremendously to the scientific understanding of chimpanzees and their behaviour. Yerkes studied chimpanzees until World War II, while Köhler concluded five years of study and published his famous Mentality of Apes in 1925 (which is coincidentally when Yerkes began his analyses), eventually concluding that "chimpanzees manifest intelligent behaviour of the general kind familiar in human beings ... a type of behaviour which counts as specifically human" (1925).
Common Chimpanzees have been known to attack humans on occasion. There have been many attacks in Uganda by chimpanzees against human children; the results are sometimes fatal for the children. Some of these attacks are presumed to be due to chimpanzees being intoxicated (from alcohol obtained from rural brewing operations) and mistaking human children for the Western Red Colobus, one of their favourite meals. The dangers of careless human interactions with chimpanzees are only aggravated by the fact that many chimpanzees perceive humans as potential rivals, and by the fact that the average chimpanzee has over 5 times the upper-body strength of a human male. As a result virtually any angered chimpanzee can easily overpower and potentially kill even a fully grown man, as shown by the attack and near death of former NASCAR driver Saint James Davis.
Chimpanzees make tools and use them to acquire foods and for social displays; they have sophisticated hunting strategies requiring cooperation, influence and rank; they are status conscious, manipulative and capable of deception; they can learn to use symbols and understand aspects of human language including some relational syntax, concepts of number and numerical sequence.  Young chimpanzees have outperformed human college students in tasks requiring remembering numbers.
Modern chimpanzees use tools, and recent research indicates that chimpanzee stone tool use dates to at least 4300 years ago. A recent study revealed the use of such advanced tools as spears, which Common Chimpanzees in Senegal sharpen with their teeth, being used to spear Senegal Bushbabies out of small holes in trees. Before the discovery of tool use in chimps, it was believed that humans were the only species to make and use tools, but several other tool-using species are now known.
Studies of language
Scientists have long been fascinated with the studies of language, as it was potentially the most uniquely human cognitive ability. To test the hypothesis of the human uniqueness of language, scientists have attempted to teach several species of great apes language. One early attempt was performed by Allen and Beatrice Gardner in the 1960s, in which they spent 51 months attempting to teach a chimpanzee, named Washoe, American Sign Language. Washoe reportedly learned 151 signs in those 51 months. Over a longer period of time, Washoe reportedly learned over 800 signs. Numerous other studies including one involving a chimpanzee named Nim Chimpsky have been conducted since with varying levels of success. There is ongoing debate among some scientists, notably Noam Chomsky and David Premack, about the great apes' ability to learn language.
Laughter in apes
Laughter might not be confined or unique to humans, despite Aristotle's observation that "only the human animal laughs". The differences between chimpanzee and human laughter may be the result of adaptations that have evolved to enable human speech. Self-awareness of one's situation such as the monkey-mirror experiments below, or the ability to identify with another's predicament (see mirror neurons), are prerequisites for laughter, so animals may be laughing in the same way that we do.
Chimpanzees, gorillas, and orangutans show laughterlike vocalizations in response to physical contact, such as wrestling, play chasing, or tickling. This is documented in wild and captive chimpanzees. Chimpanzee laughter is not readily recognizable to humans as such, because it is generated by alternating inhalations and exhalations that sound more like breathing and panting. There are instances in which non-human primates have been reported to have expressed joy. One study analysed and recorded sounds made by human babies and bonobos (also known as pygmy chimpanzees) when tickled. It found, that although the bonobo’s laugh was a higher frequency, the laugh followed a pattern similar to that of human babies and included similar facial expressions. Humans and chimpanzees share similar ticklish areas of the body, such as the armpits and belly. The enjoyment of tickling in chimpanzees does not diminish with age. 
Chimps in laboratories
As of November 2007, there were 1,300 chimpanzees housed in 10 U.S. laboratories (out of 3,000 great apes living in captivity there), either wild-caught, or acquired from circuses, animal trainers, or zoos. Most of the labs either conduct or make the chimps available for invasive research, defined as "inoculation with an infectious agent, surgery or biopsy conducted for the sake of research and not for the sake of the chimpanzee, and/or drug testing". Two federally funded laboratories use chimps: Yerkes National Primate Research Laboratory at Emory University in Atlanta, Georgia, and the Southwest National Primate Center in San Antonio, Texas. Five hundred chimps have been retired from laboratory use in the U.S. and live in sanctuaries in the U.S. or Canada.
Chimpanzees used in biomedical research tend to be used repeatedly over decades, rather than used and killed as with most laboratory animals. Some individual chimps currently in U.S. laboratories have been used in experiments for over 40 years. According to Project R&R, a campaign to release chimps held in U.S. labs — run by the New England Anti-Vivisection Society in conjunction with Jane Goodall and other primate researchers — the oldest known chimp in a U.S. lab is Wenka, who was born in a laboratory in Florida on May 21, 1954. She was removed from her mother on the day of birth to be used in a vision experiment that lasted 17 months, then sold as a pet to a family in North Carolina. She was returned to the Yerkes National Primate Research Center in 1957 when she became too big to handle. Since then, she has given birth six times, and has been used in research into alcohol use, oral contraceptives, ageing, and cognitive studies.
With the publication of the chimpanzee genome, there are reportedly plans to increase the use of chimps in labs, with some scientists arguing that the federal moratorium on breeding chimps for research should be lifted. A five-year moratorium was imposed by the U.S. National Institutes of Health (NIH) in 1996, because too many chimps had been bred for HIV research, and it has been extended annually since 2001.
Other researchers argue that chimps are unique animals and either should not be used in research, or should be treated differently. Pascal Gagneux, an evolutionary biologist and primate expert at the University of California, San Diego, argues that, given chimpanzees' sense of self, tool use, and genetic similarity to human beings, studies using chimps should follow the ethical guidelines that are used for human subjects unable to give consent. Stuart Zola, director of the Yerkes National Primate Research Laboratory, disagrees. He told National Geographic: "I don't think we should make a distinction between our obligation to treat humanely any species, whether it's a rat or a monkey or a chimpanzee. No matter how much we may wish it, chimps are not human."
An increasing number of governments are enacting a Great Ape research ban forbidding the use of chimpanzees and other great apes in research or toxicology testing. As of 2006, Austria, New Zealand, the Netherlands, Sweden, and the UK had introduced such bans.
The genus Pan is now considered to be part of the subfamily Homininae to which humans also belong. These two species are the closest living evolutionary relatives to humans. Humans shared a common ancestor with chimpanzees five to eight million years ago. Groundbreaking research by Mary-Claire King in 1973 found 99% identical DNA between human beings and chimpanzees, although research since has modified that finding to about 94% commonality, with at least some of the difference occurring in 'junk' DNA. It has even been proposed that troglodytes and paniscus belong with sapiens in the genus Homo, rather than in Pan. One argument for this is that other species have been reclassified to belong to the same genus on the basis of less genetic similarity than that between humans and chimpanzees.
A study published by Clark and Nielsen of Cornell University in the December 2003 issue of the journal Science highlights differences related to one of humankind's defining qualities — the ability to understand language and to communicate through speech. These macro-phenotypic differences, however, may owe less to physiology than might be assumed given that Homo sapiens developed modern cultural features long after the modern physiological features were in place and indeed competed averagely against other species of Homo with regard to tools, etc. for many millennia. Differences also exist in the genes for smell, in genes that regulate the metabolism of amino acids and in genes that may affect the ability to digest various proteins. See the history of hominoid taxonomy for more about the history of the classification of chimpanzees. See Human evolutionary genetics for more information on the speciation of humans and great apes.
Many human fossils have been found, but chimpanzee fossils were not described until 2005. Existing chimpanzee populations in West and Central Africa do not overlap with the major human fossil sites in East Africa. However, chimpanzee fossils have now been reported from Kenya. This would indicate that both humans and members of the Pan clade were present in the East African Rift Valley during the Middle Pleistocene.
- Template:MSW3 Groves
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- Courtney Laird. "Bonobo social spacing". Davidson College. Retrieved 2008-03-10.
- "Gene study shows three distinct groups of chimpanzees". EurekAlert. April 20 2007. Retrieved 2007-04-23. Check date values in:
- "Chimp Behavior". Jane Goodall Institute. Retrieved 2007-08-11.
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- "Chimpanzee intelligence". Indiana University. 2000-02-23. Retrieved 2008-03-24. Check date values in:
- Rowan Hooper (2007-12-03). "Chimps outperform humans at memory task". New Scientist. Retrieved 2008-03-24. Check date values in:
- Julio Mercader, Huw Barton, Jason Gillespie, Jack Harris, Steven Kuhn, Robert Tyler, Christophe Boesch (2007). "4300-year-old Chimpanzee Sites and the Origins of Percussive Stone Technology". PNAS. Feb.
- Fox, M. (2007-02-22). "Hunting chimps may change view of human evolution". Retrieved 2007-02-22. Check date values in:
- "ISU anthropologist's study is first to report chimps hunting with tools". Iowa State University News Service. 22 February, 2007. Retrieved 2007-08-11. Check date values in:
- Whipps, Heather (12 February, 2007). "Chimps Learned Tool Use Long Ago Without Human Help". LiveScience. Retrieved 2007-08-11. Check date values in:
- "Tool Use". Jane Goodall Institute. Retrieved 2007-08-11.
- "Human-like Altruism Shown In Chimpanzees". Science Daily. June 25, 2007. Retrieved 2007-08-11.
- Bradley, Brenda (1999). "Levels of Selection, Altruism, and Primate Behavior". The Quarterly Review of Biology. 74 (2): 171–194. Unknown parameter
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- "Chimpanzee lab and sanctuary map". The Humane Society of the United States. Retrieved 2008-03-24.
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- General references
- Pickrell, John. (September 24, 2002). Humans, Chimps Not as Closely Related as Thought?. National Geographic.
|Wikimedia Commons has media related to Pan.|
- Envirovet - Video clip of Ngamba Island Chimpanzee Sanctuary
- The First 100 Chimps in Research in the USA
- Chimpanzee: Wildlife summary from the African Wildlife Foundation
- Chimpanzees Make/Use Spears
- Chimpanzee Cultures Online
- Kanyawara Chimpanzee Blog from Uganda (Harvard Biological Anthropology research)
- Chimp Haven (The National Chimpanzee Sanctuary)
- Chimps as Pets (SaveTheChimps.org)
- Using Pac-Man to test cognitive reasoning in chimps
- Talking With Chimps
- Jane Goodall's Chimpanzee Central
- New Scientist 19 May 2003 - Chimps are human, gene study implies
- Did chimp and human ancestors interbreed?
- Chimp "Stone Age" Finds Are Earliest Nonhuman Ape Tools, Study Says
- Chimpanzee Facial Expression & Vocalizations
- A chimpanzee laughter sample. Goodall 1968 & Parr 2005
- Fox News: Study: Chimps Are More Evolved Than Humans
- Chimpanzees in Research: Past, Present, and Future from The State of the Animals III: 2005
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